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Coeliccia arcuata Lieftinck, 1940
(figs 5, 13, 19, 25, 31, 39, 45, 51, 57, 63, 70)
Coeliccia arcuata Lieftinck, 1940: 353-355, figs 7-8 (original description both sexes, East Kalimantan).― Lieftinck, 1954: 46 (distribution); Lieftinck, 1971: 34 (note on holotype and first described female); van Tol, 1992: 38; Orr, 2003: 39, 77; Cleary et al, 2004: 445 (East Kalimantan).
Type material.― (all RMNH): Holotype ♂, Indonesia, E. Kalimantan, Kutai, Sangkoelirang, Batu Besi, v-vi.1937, M.E. Walsh, JvT no. 2947, in brown paper triangle: “♂ ♀//Coeliccia//sp. n.”, “arcuata//Lft”, “cf//flavostriata”, “1.”, “E. Borneo, Sangkoelirang//Batu Besi//V-VI.1937 M.E. Walsh”; these all handwritten, also a red label, typed in black ink with black border “TYPES” (the ‘S’ handwritten); Paratypes: 1 ♀, in same paper triangle as holotype, same data; 2 ♀, same area and collector, Kariorang, iv.1937 and vi.1937, in paper triangle: “2 ♀”, “Coeliccia//cf. flavostriata //arcuata//Lieft.”, “sp. n.”, “♀ proth.” (this circled), “E. Borneo//Sangkoelirang”, “PARATYPES”, “Kariorang”, “IV.1937”, “id., VI.1937” and “M.E. Walsh”; 1 ♂, same area and collector, vi.1937, Babi Djoeton.
Other material.― Sabah (in RMNH except as noted): 1 ♂, N Sabah, Mt Kinabalu, Poring Hot Springs, small stream near trail to Langanan waterfall, 600-700 m, 29.iv.2005, in coll. Dow; 1 ♂ (JvT no. 8085), 16.iii.1987, E. Sabah, 60 km W. of Lahad Datu, Danum Valley, near Sungai Segama, brooklet crossing west trail, 170m, J. van Tol; 8 ♂ (JvT nos. 8078-8083, 8086-8087), 19.iii.1987, same area, brooklet above artificial pond near field centre, 1 misidentified as C. campioni, J. van Tol; 1 ♂ (JvT no. 8088), 24.iii.1987, same area, Sungai Palum Tambon, 160-180 m, J. van Tol; 2 ♂ (1 in pieces), 23-28.iv.1994, same site, 150-160 m, M. Hämäläinen; 1 ♂, 9-18.xi.1958, E. Sabah, Tawau, Kalabkan river, T.C. Maa. Indonesia, Kalimantan (all RMNH): 2 ♂, 12.xi.1996, S. Kalimantan, Banjarbaru, Aranio distr., 5 km SE of Belangian village, Riam Kanan Lake, Gunung Pisang primary forest, Sungai Kuinam, M. Bedjanič.
Remarks.― C. arcuata is easily distinguished from all other known species except C. species A by the form of the male paraprocts in ventral view (fig. 63) and the posterior pronotal lobe in the female (figs 5, 13, 19). It is most readily distinguished from C. sp. A by the position and size of the tooth on the cerci, and by the paraprocts seen in profile (figs 51, 56).
Lieftinck (1940: 353) lists two males and three females in the type series, but only the holotype and three female paratypes are marked as such in RMNH. However an additional male with data coinciding with that of the type series, collected at “Babi Djoeton”, a location listed by Lieftinck for the type series, is present and must be the additional male paratype; it is listed as such above.
Lieftinck’s (1940) description of both sexes of arcuata requires few additions, however it does not give the colour of the antehumeral stripes of male arcuata; in the holotype and some other males they are yellow, but others have blue antehumeral stripes; males with different coloured antehumeral stripes can occur within single populations. The antehumeral stripes of the female paratypes are blue. It is likely the colour of the antehumeral stripes is correlated with the maturity of the individual. There are blue markings on the dorsum of the axillaries in the male, and on the metascutum.
The prothorax of female arcuata (figs 5, 19) has large notopleural projections, but no anterior lobe processes or median lobe projections. The raised part of each mesostigmal plate is separated from the posterior pronotal lobe by nearly its own width.
In the type series the dorsal surface of S2 has no pale markings, but in all other material studied a blue, oval to rectangular marking is present on the basal two-thirds. The extent of the blue markings on S9-10 also varies, in some specimens the entire dorsum is blue, in others (including the holotype and male paratype) these markings are reduced to a large dorsal mark on S9 and a much smaller mid dorsal spot on S10.
The measurements of material studied fall within the following ranges: ♂: abdomen without appendages 33-38.75 mm, Hw 19.5-24 mm; ♀: abdomen without cerci or ovipositor 30-33 mm, Hw 21-22 mm.
Distribution and habitat.― Northern and eastern Sabah from Poring Hot Springs southwards, southern and eastern Kalimantan (fig. 70). Small streams and springs in lowland mixed dipterocarp forest.
