Zoologische Mededelingen, 85 (April 2011)D. van den Tooren: New records of olpiid pseudoscorpions (Pseudoscorpiones: Olpiidae) from the Caribbean area and Surinam, with descriptions of four new species of the genera Pachyolpium Beier, Tricholpium gen. nov. and Heterohorus gen. nov.

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Pachyolpium dispersum spec. nov.

(figs 2 a-f, 3a-d, 4a-b)

Material examined.― Holotype: ♀ (1281), Tortuga: # 173, South-western Tortuga, 1.viii.1936. Altitude: 20 m; coral-limestone; shrubs and small trees with much Condalia and cacti; under flat stones with some plant-decay; paratypes: 3 ♀♀ (852-853, 856), Rancho Grande (Venezuela): # 555, Parque Nac. Pittier, 1½ km E of Estacíón, 17.iii.1955. Altitude: 1200 m; schists, “selva nublade”, plant decay. Allotype: ♂ (854) and paratype: ♂ (855); paratypes nymphs: 2 tritonymphs (857-858), 1 deutonymph (859), and 3 proto-nymphs (864, 870, 883).

Diagnosis.― The new species differs from the related Pachyolpium amplum (= “Olpiolum” amplum) Hoff (1964: 24-25; only one male known from Jamaica) by the position of the trichobothria on the chelal fingers and the position of the nodus ramosus on chelal movable finger.

Description.― Female: description based on the holotype and three paratypes. Carapace practically smoothly dark reddish orange, tergites, chelicerae and legs pale yellowish orange, segments of pedipalps unequal in colour: trochanter yellowish orange, femur and tibia bright orange, and chela deep reddish orange. Body L 2.95 (2.77-3.17).

Carapace: 0.80 (0.79-0.80) / greatest width 0.79 (0.76-0.84), L/W 1.01 (0.94-1.05); eyes: 2 pairs, well developed, anterior eye half its diameter from posterior eye and about 1.4 times its own diameter from the anterior margin of the carapace; anterior margin with 6 setae, on posterior margin no more than 4 setae perceptible.

Abdomen: chaetotaxy tergites I-X: 5.5(4-8): 6.0(4-8): 8.5(8-10): 13.0(8-12): 11.5(8-16): 11.25(10-15): 11.75(10-15): 11.75(10-15): 1T7-10T1 (3): 1T3-8T1: chaetotaxy sternites IV-XI: 7.3(6-8): 13.7(12-16): 14.0(11-16): 12.3(12-13): 11.0(10-13): 10.7(10-12): 9.0(6-11): 2T4T2 (1).

Chelicera (figs 3a-b): 0.355 (0.31-0.35) / 0.19 (0.18-0.20), L/W 1.78 (1.70-1.85), and 0.42 (0.39-0.44) × carapace L; movable finger L 0.24 (0.20-0.28); galea L 0.09 (0.06-0.105), straight stem with 3 terminal curved rami of average length; inner margin of fixed finger with about 5 very flat non-sclerotic retroconical teeth ranging from distal end up to half-way finger length, and most distal a small sclerotic denticle, situated on the inner margin of the sclerotic apical tooth; movable finger with prominent sharp subapical lobe; serrula exterior with 23.75 (23-25) ligulate blades, flagellum of 3 spine-like setae: anterior most one longest (L: 0.09-0.10) with about 7 small spinules along distal end of anterior margin, middle one (L: 0.06-0.065) with spinules along entire anterior margin and posterior most one (L: 0.05-0.06) with about 4 small spinules in distal end of anterior margin; chaetotaxy: 5 setae on exterior surface of hand and 1 sub-apical seta (galeal seta) on exterior surface of movable finger, somewhat shorter than galea. Lamina exterior present.

Pedipalp, practically smooth (fig. 2a-b): trochanter 0.44 (0.39-0.46) / 0.27 (0.27-0.28), L/W 1.615 (1.45-1.72); femur 0.76 (0.73-0.78) / 0.26 (0.25-0.265), L/W 2.95 (2.77-3.09); patella 0.795 (0.79-0.80) / 0.335 (0.32-0.35), L/W 2.37 (2.26-2.46); femur L/patella L 0.96 (0.92-0.98); chela without pedicel 1.35 (1.30-1.37) / 0.52 (0.49-0.55), L/W 2.60 ( 2.48-2.67); chela with pedicel 1.445 (1.40-1.47) / 0.52 (0.49-0.55), L/W 2.785 (2.65-2.87); hand without pedicel 0.725 (0.68-0.75) / 0.52 (0.49-0.55), L/W 1.40 (1.36-1.43); hand with pedicel 0.83 (0.78-0.86) / 0.52 (0.49-0.55), L/W 1.60 (1.56-1.62); movable finger L 0.70 (0.68-0.71), and 0.965 (0.92-1.03) × L hand without pedicel, 0.84 (0.81-0.89) × L hand with pedicel; sb on dorsum palpal femur 0.20 (0.18-0.21) from exterior key point of femoral base, i.e. palpal femur L 3.88 (3.58-4.38) × distance from sb to exterior key point of femoral base.

Number of marginal teeth in fixed finger (3) 51 (50-51), in movable finger (3) 56 (50-60). Marginal teeth in fixed - and movable finger of different shape, except the small triangular and cuspidated teeth at the distal end of the teeth row; in fixed finger ranging from broader and flatter to very flat and undeveloped teeth at the basal end of row; in movable finger from slightly retroconical to more rounded smaller teeth, and extending 4 teeth farther to the finger base than in fixed finger.

Distance trichobothria and nodus ramosus from base apical tooth on outside curve in fixed finger: et 0.085 (0.08-009), est 0.38 (0.36-9.39), esb 0.60 (0.55-0.64), eb 0.63 (0.585-0.67), it 0.29 (0.28-0.30), ist 0.49 (0.47-0.51), isb 0.55 (0.515-0.58), ib 0.57 (0.56-0.58), and nodus ramosus 0.12 (0.10-0.14), i.e. et a little distal to the level of nodus ramosus and located in distal seventh of finger L (finger L arbitrarily considered equal to movable finger L), it and est close to median point finger, it a little distal, est a little proximal to that point, remaining trichobothria within basal third of finger L.

Distance trichobothria and nodus ramosus from base apical tooth on outside curve in movable finger: t 0.26 (0.24-0.27), st 0.50 (0.48-0.505), sb 0.585 (0.57-0.60), b 0.62 (0.60-0.64), and nodus ramosus 0.13 (0.08-0.15), i.e. t between halfway and one third finger L from fingertip, nodus ramosus halfway level t and finger tip, st about one fifth finger L proximal to midpoint finger, and sb and b within basal seventh of finger L.

First leg (fig. 3c): trochanter 0.21 (0.20-0.22) / 0.16 (0.16-0.17), L/D 1.325 (1.21-1.44); femur 0.38 (0.36-0.40) / 0.12 (0.12-0.13), L/D 3.04 (2.94-3.09); patella 0.22 / 0.13 (0.12-0.13), L/D 1.73 (1.68-1.79); femur L 1.73 (1.68-1.79) × patella L; tibia 0.31 (0.30-0.315) / 0.09, L/D 3.43 (3.34-3.60); metatarsus 0.235 (0.21-0.245) / 0.07 (0.06-0.07), L/D 3.54 (3.43-3.73); tarsus 0.175 (0.17-0.18) / 0.05 (0.05-0.55), L/D 3.22 (3.17-3.33); L claws 0.055 (0.05-0.06); arolium L 0.07 (3), i.e. 1.20 (1.15-1.23) × L claws (3).

Fourth leg (fig. 3d): trochanter 0.34 / 0.19 (0.175-0.19), L/D 1.82 (1.71-1.95); femur 0.25 (0.24-0.26) / 0.165 (0.16-0.17), L/D 1.50 (1.45-1.52); patella (3) 0.62 (0.61-0.63) / 0.26 (0.255-0.26), L/D 2.44 (2.40-2.47); femur + patella 0.755 (0.74-0.77) / 0.26 (0.255-0.26) (3), L/D (3) 2.97 (2.99-3.0); tibia 0.545 (0.53-0.56) / 0.13 (0.125-0.13) (3), L/D (3) 4.30 (4.22-4.38); metatarsus 0.34 (0.31-0.36) / 0.09, L/D 3.80 (3.33-4.15); tarsus 0.24 (0.23-0.255) / 0.69 (0.065-0.07), L/D 3.50 (3.40-3.57); L claws 0.075 (0.07-0.08); arolium 0.10, i.e. 1.34 (1.27-1.40) × L claws.

Chaetotaxy of metatarsus of fourth leg: exterior lateral T+2 (T near proximal end), dorsal 3, interior lateral (3-4) × 2 (2) and (1-3) × 2 + 1 (2), ventral 3.

Chaetotaxy and internal structure of female genital area: anterior operculum on face with 3 setae in a practically straight line forward to the centre on each side of median line; near the anterior side of the segment a barely perceptible medial cluster of 10 or 15 very small setae on each side of median line; a moderate medial oval cribriform plate expanding somewhat sideward and possibly also with small lateral plates.

Discussion (includes also data obtained from examined male specimens out of additional material, which has been described after this).― Pachyolpium dispersum spec. nov. is distinguished from P. amplum Hoff, 1964, from Jamaica by the position of the trichobothrium est on chelal fixed finger: about twice in male, 1.2 × in female, farther from the level of ist than to the level of it (versus est distinctly closer to the level of ist than to the level of it); the position of the nodus ramosus on chelal movable finger: about 1.2 × in male, 1.0 × in female (versus twice) as far from finger tip as from the level of t; and st about 2.85 × both in male and female (versus somewhat more than 1.5 ×) farther from the level of t than from that of sb.

Male.― Description based on the allotype (854) and 1 paratype (855) (figs. 4a-b). Ratios and counts expressed as ranges. Colour about similar to that of female; body, chelicerae and legs on the whole much slimmer than in ♀. Body L 2.42/2.45.

Carapace: 0.71/0.73/greatest W 0.63/0.64, L/W 1.125/1.14, eyes 2 pairs, well developed, eyes of each pair very close together, anterior eye about its own diameter from anterior margin of carapace; anterior margin with 6/6 setae, posterior margin with 4/4 setae.

Abdomen: chaetotaxy of tergites I-X: 4/4:4/4: 9/9: 11/10: 11/13: 11/14: 11/12: 12/13: 12/12: 1T6T1/1T8T1; chaetotaxy of sternites IV-X: 9/10: 13/17: 13/17: 13/15: 13/13: 11/11: 9/10.

Chelicera: 0.29/0.30 / 0.16/0.17, L.W 1.74/1.84, and 0.41/0.41 × carapace L; movable finger L: 0.22/0.23; galea L 0.05/0.05, straight stem with 3 short curved rami; inner margin of fixed finger as in female; movable finger with blunt conical subapical lobe joined with 2 tiny marginal triangular thorns; serrula exterior with 22/23 ligulate blades, flagellum of 3 spine-like setae, anterior most one longest (L: 0.13), with small spinules in distal half of anterior margin, middle one (L: 0.11), with small spinules in distal third of anterior margin, and most posterior one (L: 0.095), with some small spinules in distal quarter of anterior margin; chaetotaxy as in female. Lamina exterior present.

Pedipalp: trochanter 0.38/0.39 / 0.225/0.23, L/W 1.67/1.72; femur 0.65/0.70 / 0.22/0.22, L/W 2.96/3.20; patella 0.69/0.71 / 0.275/0.29, L/W 2.45/2.51; femur L/patella L 0.95/0.985; chela without pedicel 1.13/1.15 / 0.40/0.405, L/W 2.83/2.84; chela with pedicel (fig. 4a) 1.21/1.23 / 0.40/0.405, L/W 3.02/3.05; hand without pedical 0.59/0.60 / 0.40/0.41, L/W 1.47/1.50; hand with pedicel 0.68/0.68 / 0.40/0.41, L.W 1.67/1.72; movable finger L 0.58/0.63, and 0.98/1.04 × L. hand without pedical, 0.85/0.92 × L hand with pedicel; sb on dorsal palpal femur 0.175/0.19 from exterior key point of femoral base, i.e. palpal femur L 3.68/3.71 × distance from sb to exterior key point of femoral base.

Number of marginal teeth in fixed finger 48/52, in movable finger 55/61. Shape of marginal teeth in both fingers about as in female. Distance trichobothria and nodus ramosus from base apical tooth on outside curve in fixed finger: et 0.07/0.08, est 0.30/0.34, esb 0.49/0.55. eb 0.51/0.57, it 0.25/0.27, ist 0.43/0.45, isb 0.475/0.51, ib 0.49/0.52, and nodus ramosus 0.10/0.11, i.e. position of trichobothria and nodus ramosus about as in female. Distance trichobothria and nodus ramosus from base apical tooth on outside curve in movable finger: t 0.20/0.22, st 0.39/0.43, sb 0.45/0.51, b 0.49/0.55, and nodus ramosus 0.11/0.12, i.e. position of trichobothria and nodus ramosus about as in female.

First leg: trochanter 0.19/0.20 / 0.14/0.14, L/D 1.37/1.38; femur 0.34/0.34 / 0.11/0.11, L/D 3.02/3.12; patella 0.19/0.19 / 0.11/0.11, L/D 1.72/1.72; femur L 1.76/1.77 × patella L; tibia 0.27/0.28 / 0.08/0.09, LD 3.265/3.56; metatarsus 0.22/0.22 / 0.06/0.06, L/D 3.47/3.48; tarsus 0.16/0.165 / 0.05/0.05, L/D 3.07/3.17; L claws 0.05/0.05; arolium L 0.06 (1) (stem retracted).

Fourth leg: trochanter 0.29/0.31 / 0.15/0.17, L/D 1.80/1.94; femur 0.21/0.23 / 0.13/0.14, L/D 1.57/1.75; patella 0.49/0.54 / 0.24/0.25, L/D 1.94/2.25; femur + patella 0.66/0.69 / 0.24/0.25, L/D 2.71/2.72; tibia 0.48/0.49 / 0.12/0.12, L/D 3.93/3.97; metatarsus 0.30/0.31 / 0.08/0.08, L/D 3.73/3.74; tarsus 0.22/0.23 / 0.06/0.07, L/D 3.53/3.60; L claws 0.065/0.07; arolium 0.08/0.09, and 1.21/1.325 × L claws.

Chaetotaxy of metatarsus of fourth leg: exterior lateral T+2/T+2 (T close to proximal end), dorsal 2/3, interior lateral 4×2/4×2, ventral 3/3.

Chaetotaxy of male genital area (number of setae given for both sides of median line) (fig. 4b): anterior operculum: 4-3/4-3, posterior margin of anterior lip: 2-1 (1), on face: 3-4 (1), anterior margin of posterior lip 2-2 (1), posterior margin of posterior lip: 2-2/2-2, posterior operculum: 2-3/2-2.

Tritonymph. Description based on 2 specimens (857-858) from Rancho Grande (Venezuela). Ratios and counts expressed as ranges. Body L 1.97/2.06.

Carapace: 0.61/0.63 / greatest W 0.46/0.46, L/W 1.32/1.36; eyes 2 pairs; anterior margin with 6 setae, posterior margin with 4 setae.

Abdomen: chaetotaxy sternites I-X: 4/4: 6/4: 9/8: 8/8: 9/8: 9/10: 9/9: 7/12: 1T7T1 (1): 1T4T1 (1); chaetotaxy sternites IV-X: 8/7: 8/10: 10/10: 10/11: 8/10: 8/8: 8/8.

Chelicera: 0.25/0.26 / 0.14/0.15, L/W 1.71/1.81, and 0.41/0.41 × carapace L; movable finger L 0.17/0.20; galea L 0.07/0.08 with 3 short terminal rami; inner margin of fixed finger with 4 very flat retroconical teeth associated with apical tooth; movable finger with sharp conical subapical lobe; serrula exterior of 21/21 ligulate plates; flagellum of 3 spine-like setae:, anterior most one longest (L.: 0.07/0.07), L middle one: 0.045/0.005, and L posterior most one: 0.04/0.04, on anterior margin of all 3 setae small spiny denticles perceptible. Chaetotaxy complete: 5 setae on exterior surface of hand and 1 on movable finger.

Pedipalp: trochanter 0.315/0.33 / 0.18/0.20, L/W 1.67/1.71; femur 0.49/0.53 / 0.175/0.18, L.W 2.80/2.93; patella 0.53/0.54 / 0.23/0.24, L.W 2.30/2.35; femur L 0.92/0.97 × patella L; chela without pedicel 0.92/0.97 / 0.34/0.35, L/W 2.68/2.73; chela with pedicel 1.01/1.03 / 0.34/0.35, L/W 2.91/2.96; hand without pedicel 0.48/0.50 / 0.34/0.35, L/W 1.41/1.42; hand with pedicel 0.55/0.61 / 0.34/0.35, L/W 1.62/1.73; movable finger L: 0.485/0.50, and 1.0/1.01 × L hand without pedicel, and 0.82/0.88 × L hand with pedicel; sb on dorsum femur 0.15/0.15 from exterior key point of femoral base.

Number of marginal teeth in fixed finger 40 (1), in movable finger 45 (1). Distance trichobothria and nodus ramosus from base apical tooth on outside curve in fixed finger: et 0.07/0.07, est 0.26/0.27, esb 0.41/0.42, eb 0.44/0.45, it 0.23/0.23, ist 0.36/0.37, isb lacking, ib 0.41/0.42 and nodus ramosus 0.09/0.09, i.e. et and nodus ramosus within ⅕ finger L (arbitrarily considered equal to movable finger L), level nodus ramosus a little proximal to that of et, it about median point finger, est proximal to that point, ist on a level of about ¼ finger L from base, ib, b and esb in a still more basic position.

Distance trichobothria and nodus ramosus in movale finger: t 0.19/0.20, st 0.35/0.37, sb lacking, b 0.42/0.44, and nodus ramosus 0.09, i.e. t on about a level of ⅖ finger L from finger tip, nodus ramosus about halfway t and finger tip, st about ¼ finger L from base, and b about halfway the level of st and finger base.

First leg: trochanter 0.16/0.17 / 0.11/0.12.5, L/D 1.34/1.38; femur 0.26/0.27 / 0.09/0.10, L/D 2.69/2.86; patella 0.16/0.16 / 0.09/0.10. L/D 1.63/1.70; femur L 1.67/1.74 × patella L; tibia 0.20/0.22 / 0.07/0.07, L/D 2.95/3.01; metatarsus 0.15/0.17 / 0.05/0.06, L/D 2.82/2.92; tarsus 0.14/0.15 / 0.05, L/D 3.05/3.15; L claws 0.035/0.04; L arolium 0.04/0.05, and 1.12/1.265 × L claws.

Fourth leg (1): trochanter 0.23/0.135, L.D 1.69; femur 0.18/0.135, L/D 1.36; patella 0.44/0.20, L/D 2.18; femur + patella 0.54/0.20, L/D 2.68; tibia 0.385/0.10, L/D 3.93; metatarsus 0.24/0.075, L/D 3.07; tarsus 0.18/0.05, L/D 3.47; L claws 0.035; L arolium 0.04, and 1.25 × L claws.

Chaetotaxy of metatarsus of fourth leg: exterior lateral: T+2, dorsal: 2, interior lateral: 3×2, ventral: 2.

Deutonymph. Description based on the specimen (859) from Rancho Grande (Venezuela).

Body L 1.70. Carapace: 0.52/greatest W 0.38, L/W 1.37, eyes 2 pairs, position as in tritonymph, anterior margin of carapace with 6 setae, posterior margin with 4 setae.

Abdomen: chaetotaxy of tergites IV-XI: 6: 8: 10: 10: 10: 10: 8: 1T3T1; chaetotaxy of sternites IV-XI: 8: 8: 8: 9: 8: 8: 7: 1T3T1.

Chelicera: 0.20/0.12. L/W 1.70, and 0.38 × carapace L, movable finger L 0.15, galea L 0.06, with 3 short curved terminal rami; inner margin of fixed finger with 4 very flat retroconical teeth associated with apical tooth; movable finger with prominent sharp conical subapical lobe; serrula exterior with 19 ligulate blades; flagellum of 3 spine-like setae, anterior most one longest (L.: 0.06), L middle one: 0.035 and L posterior most one: 0.03; on anterior margin of first 2 setae small spinules in distal third. Chaetotaxy as before, all setae are present.

Pedipalp: trochanter 0.25/0.15, L/W 1.71; femur 0.41/0.14, L/W 2.97; patella 0.39/0.175, L/W 2.255; femur L 1.04 × patella L; chela without pedicel 0.71/0.245, L/W 2.89; chela with pedicel 0.75/0.245, L.W 3.09; hand without pedicel 0.36/245, L/W 1.46; hand with pedicel 0.39/0.245, L/W 1.61; movable finger L 0.35, and 0.97 × L hand without pedical, and 0.88 × L hand with pedicel; sb on dorsum femur 0.12 from exterior key point of femoral base, i.e. palpal femur L 3.42 × distance from sb to exterior key point of femoral base.

Number of marginal teeth in fixed finger up to about 35, in movable finger up to about 40 marginal teeth. Distance trichobothria and nodus ramosus from base apical tooth on outside curve in fixed finger: et 0.09, est 0.22, esb lacking, eb 0.35, it 0.20, ist 0.25, isb lacking, ib 0.33, and nodus ramosus 0.07, i.e. nodus ramosus at about distal sixth, it just distal to median point and ist, ib and eb within basal fifth of finger (arbitrarily considered equal to movable finger L). Distance trichobothria and nodus ramosus from base apical tooth on outside curve in movable finger: t 0.175, st 0.32, sb and b lacking, and nodus ramosus 0.07, i.e. nodus ramosus at about distal sixth of finger, t distal to median point finger, and st at about basal quarter of finger.

First leg: trochanter 0.13/0.10, L/D 1.32; femur 0.20/0.065, L/D 3.07; patella 0.12/0.07, L/D 1.595; femur L 1.72 × patella L; tibia 0.16/0.06, L/D 2.88; metatarsus 0.115/0.05, L/D 2.38; tarsus 0.12/0.04, L/D 3.0; L claws 0.035, L arolium 0.04, and 1.20 × L claws.

Fourth leg: trochanter 0.18/1.05, L/D 1.75; femur 0.13/0.09, L/D 1.38; patella 0.32/0.135, L/D 2.35; femur + patella 0.39/0.135, L/D 2.92; tibia 0.29/0.08, L/D 3.67; metatarsus 0.17/0.06, L/D 2.83; tarsus 0.15/0.04, L/D 3.44, L claws 0.05; L arolium 0.06, and 1.24 × L claws.

Chaetotaxy of metatarsus of fourth leg: exterior lateral: T+1, dorsal: 1, interior lateral: 2x2, ventral: 1.

Protonymph. Description based on 3 specimens (864, 870, 883) from Rancho Grande, Venezuela. Body L 1.24 (1.16-1.305).

Carapace: 0.45 (0.45-0.46) / greatest W 0.34 (0.32-0.36), L/W 1.335 (1.29-1.38); eyes 2 pairs, front eye of each pair about 2 times its own diameter from anterior margin; anterior carapacal margin with 4 (2) or 2 (1) marginal setae, posterior margin with 2 (1) or 4 (2) marginal setae.

Abdomen: middle tergites and middle sternites with 4 marginal setae.

Chelicera: 0.17 (0.16-0.17) / 0.10 (0.095-0.10) / 1.71 (1.65-1.74), and 0.37 × carapace L; movable finger L 0.12 (0.115-0.12); galea L 0.05, with 3 short curved terminal rami; teeth on inner margin of fixed finger and subapical lobe on movable finger about as in deutonymph; serrula exterior with 15 ligulate blades; flagellum of 3 spines, anteriormost one longest (L: 0.03/0.05 (2)), L middle one: 0.03 (2), and L posterior one: 0.02 (2). Chaetotaxy: number of setae on exterior surface of cheliceral hand incomplete, lacking: galeal seta and sub-basal seta.

Pedipalp: trochanter 0.19 (0.19-0.20) / 0.11 (0.105-0.12), L/W 1.73 (1.62-1.85); femur 0.30 (0.29-0.30) / 0.105 (0.10-0.11), L/W 2.90 (2.82-3.0); patella 0.30 (0.29-0.315) / 0.13 (0.12-0.13), L/W 2.38 (2.32-2.41); femur L 0.99 (0.96-1.02) × L patella; chela without pedicel 0.56 (0.53-0.57) / 0.18 (0.16-0.185), L/W 3.13 (3.01-3,26); chela with pedicel 0.59 (0.56-0.615) / 0.18 (0.16-0.19), L/W 3.34 (3.185-3.48); hand without pedicel 0.28 (0.27-0.30) / 0.18 (0.16-0.19), L/W 1.59 (1.435-1.71); hand with pedicel 0.325 (0.315-0.34) / 0.18 (0.16-0.19), L/W 1.83 (1.68-1.95); movable finger L 0.30 (0.27-0.32), and 1.06 (0.98-1.20) × L hand without pedicel, and 0.92 (0.87-1.04) × hand with pedicel; sb on dorsum femur 0.09 from exterior key point of femoral base, i.e. palpal femur L 3.18 (3.0-3.22) × distance from sb to exterior key point of femoral base.

Number of marginal teeth in fixed finger 28 (1), in movable finger 24 (1) marginal teeth. Distance trichobothria and nodus ramosus from base apical tooth on outside curve in fixed finger: et 0.12 (0.11-0.13), est lacking, esb lacking, eb 0.28 (0.26-0.30), it lacking, ist 0.25 (0.24-0.26), isb lacking, ib lacking and nodus ramosus in a distal position 0.06 (0.06-0.07), i.e. nodus ramosus at about one fifth finger L from finger tip (finger L arbitrarily considered equal to movable finger L), et a little distal to the level of nodus ramosus and eb in a basal position. Distance trichobothria and nodus ramosus from base apical tooth on outside curve in movable finger: t 0.18 (0.17-0.20), st lacking, sb lacking, b lacking, and nodus ramosus 0.06 (0.06-0.07), i.e. nodus ramosus in same position as in fixed finger and t about three-fifth finger L from finger tip.

First leg: trochanter (2) 0.09-0.105 / 0.07-0.08, L.D 1.22/1.33; femur 0.14 (0.13-0.15) / 0.05 (0.05-0.06), L/D 2.46 (2.40-2.50); patella 0.09 (0.06-0.07), L/D 1.49 (1.42-1.54); femur L 1.52 (1.43-1.61) × patella L; tibia 0.12 (0.11-0.12) / 0.05, L/D 2.32 (2.155-2.47); metatarsus 0.08 (0.08-0.09) / 0.04 (0.04-0.05), L.D 2.0; tarsus 0.10 (0.095-0.11) / 0.04 (0.03-0.04), L/D 2.71 (2.45-2.97); L claws 0.03; L arolium 0.03 (0.03-0.04), and 1.09 (0.92-1.22) × L claws.

Fourth leg: trochanter (2) 0.14-0.15 / 0.08, L/D 1.78-1.94; femur 0.11 (0.10-0.12) / 0.08 (0.08-0.09), L.D 1.34 (1.33-1.36); patella 0.24 (0.22-0.25) / 0.11 (0.10-0.11), L/D 2.28 (2.27-2.29); femur + patella 0.30 (0.28-0.32) / 0.11 (0.10-0.11), L/D 2.89 (2.85-2.94); tibia 0.22 (0.21-0.23) / 0.06 (0.06-0.65), L/D 3.55 (3.51-3.57); metatarsus 0.135 (0.125-0.14) / 0.05, L/D 2.74 (2.60-2.90); tarsus 0.13 (0.12-0.135) / 0.04, L/D 3.21 (3.08-3.32); L claws 0.04; L arolium (2) 0.04-0.05, and 0.99-1.23 × L claws.

Chaetotaxy of metatarsus of fourth leg: exterior lateral: T+1, dorsal: 1, interior lateral: 2×1, ventral: 1.

Etymology.― the specific epitheton (dispersum) indicates that the description of the new species is based on specimens originated from different localities (dispersus (Latin for scattered).

Remarks.― It appears that in all three nymphal stages the trichobothria isb and sb on respectively palpal fixed- and movable finger are lacking, same for esb and st in proto- and deutonymph, and further for est, it, ib and b in protonymph. According to Harvey (1992: 1421, figs 172-175) a statement valid for all species of the superfamily Olpioidea Banks, stat. nov. The setae on the exterior surface of the chelicera (5 on hand, 1 on movable finger) are present in trito- as well in deutonymph, however in proto-nymph sub-, basal- and galeal setae are absent.

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