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Key to species of genus Vietorogas nov.
1. Vein 1-CU1 of fore wing about as long as vein m-cu and first discal cell weakly narrowed distally (fig. 11); vein 1-M of hind wing slightly shorter than 1r-m; mesoscutum irregularly rugose posteriorly (fig. 15); precoxal sulcus largely transversely rugose (fig. 16); propodeum reddish yellow and largely rugose with a strong median carina anteriorly; propodeal spiracle close to anterior border of propodeum (fig. 12); vein 1-SR of fore wing comparatively long (fig. 11); propodeal spiracle medium-sized (fig. 12) → V. abditivus Long & van Achterberg, spec. nov.
- Vein 1-CU1 of fore wing distinctly longer than vein m-cu and first discal cell distinctly narrowed distally (fig. 4); vein 1-M of hind wing as long as 1r-m; mesoscutum finely rugose posteriorly; precoxal sulcus finely and sparsely rugose; propodeum dark brown, but pale yellow medially and completely rugose-granulate with a less strong median carina anteriorly; propodeal spiracle comparatively far from anterior border of propodeum (fig. 8); vein 1-SR of fore wing short (fig. 4); propodeal spiracle small (fig. 8) → V. bachma Long, spec. nov.
Cornutorogas aurantius van Achterberg, spec. nov.
next sectionMaterial.— Holotype, ♀ (RMNH), “S. Vietnam: Dông Nai, Cát Tiên N.P., Mal. traps, c. 100 m, 9.iv.-13.v.2007, Mai Phu Quy & Nguyen Thanh Manh, RMNH’07”. Paratype: 1 ♂ (IEBR), “S. Vietnam: Dak Lak, Chu Yang Sin N.P., n[ea]r dam, 740-940 m, 1-10.vi.2007, Mal. traps, C. v. Achterberg & R. de Vries, RMNH’07”.
Holotype, ♀, length of body 5.7 mm, length of fore wing 4.0 mm.
Head.— Antenna with 60+ segments and 2.5 times as long as fore wing, third segment 1.2 times as long as fourth segment, length of third, fourth and penultimate segments 5.2, 4.2 and 1.8 times their width, respectively; scapus longer dorsally than ventrally in lateral view; scapus 1.6 times longer than apically wide; occipital carina strongly developed (fig. 50); eye large and emarginate, resulting in a subtriangular head in anterior view (fig. 49); in anterior view height of eye 1.7 times its width; eye 1.3 times longer than width of face; tentorial pits large, distance between tentorial pits 1.5 times distance from pit to eye (fig. 49); face 1.2 times wider than height of face and clypeus combined; in lateral view width of eye 5.4 times central width of temple (fig. 50), in dorsal view 4.8 times temple; POL:Od:OOL = 5:6:8; face largely smooth, sparsely punctulate and setose; vertex and temple sparsely finely punctate; length of malar space 1.2 times basal width of mandible.
Mesosoma.— Length of mesosoma (without propodeal horn) 1.5 times its height; pronotum comparatively robust and with large pronope (fig. 52) and in dorsal-lateral view hardly convex medio-posteriorly (fig. 53); side of pronotum rather slender (fig. 51); prepectal carina lamelliform; precoxal sulcus rather deep and strongly carinate, absent near middle coxa; dorsally mesopleuron coarsely crenulate and below it and ventrally spaced punctate; mesoscutum setose and punctulate; notauli rather narrow, deep and coarsely carinate; median lobe of mesoscutum with rather short and largely shallow (except posterior pit) groove posteriorly; axillar lobe rather wide and obtuse (fig. 56); scutellum sparsely punctulate; propodeum coarsely vermiculate-rugose; lateral horn of propodeum 0.8 times as long as first metasomal tergite.
Wings.— Fore wing: pterostigma 4 times as long as wide; vein r from middle of pterostigma; length of vein r:2-SR:3-SR:SR1 = 10:17:19:60; length of vein 1-CU1:cu-a:2-CU1:3-CU1 = 5:10:43:7; second submarginal cell rather short, parallel-sided, 1.1 times longer than vein r-m. Hind wing: M+CU:1-M: 1r-m = 40:53:16.
Legs.— Hind coxa rather densely punctate; length of fore femur:tibia:tarsus:basitarsus = 50:57:62:25; length of femur, tibia and basitarsus of hind leg 5.6, 9.0 and 10.4 times their width, respectively; both hind tibial spurs 0.2 times as long as basitarsus.
Metasoma.— First tergite 1.2 times as long as wide apically, basally hardly widened beside dorsope, its surface coarsely longitudinally costate; dorsal carinae remain separated; second tergite with small but distinct semicircular medio-basal area and complete median carina continuing up to 0.9 of third tergite; second tergite 1.4 times as long as third tergite; second tergite sublongitudinally costate, third and fourth tergites coarsely longitudinally rugose-striate and between striae punctate; fifth tergite densely coarsely discretely punctate but laterally rugose; length of ovipositor sheath 0.5 times basitarsus and 0.13 times fore wing, its apical half somewhat widened.
Colour.— Head black; mandible, palpi, pronotum (except posteriorly), scapus and pedicellus (except dark outer streak), fore and middle coxae, trochanters and trochantelli, humeral plate, propodeal horn, apex of fore and middle femora, base and apex of all femora, base and apex of hind basitarsus and second-fifth tarsal segments, ivory; 3rd-8th antennal segments brownish-yellow, 9th-28th and apical segments blackish and 29th-53rd segments white; mesoscutum dark orange red but laterally narrowly darkened; tegulum, pronotum posteriorly and mesopleuron anteriorly blackish; hind trochantellus, apex of hind femur, remainder of hind tibia and of all basitarsi dark brown; remainder of mesopleuron and mesosternum orange brown; remainder of mesosoma (including axillae and scutellum) and remainder of legs brownish-yellow; metasoma pale yellowish ventrally, largely orange brown dorsally, but all tergites (except second) partly more or less darkened; ovipositor sheath, most veins and pterostigma (except narrowly pale base) dark brown; base of pterostigma and parastigma largely pale yellowish; wing membrane slightly infuscate, parastigma yellow.
Male.— Similar to female but head normal, not subtriangular, antenna with 56 segments and 29th-34th antennal segments white and 35th segment greyish; length of body 5.6 mm and length of fore wing 4.5 mm; apex of hind coxa partly blackish; hind tibial spurs of ♂ slender and acute apically; third-fifth tergites largely blackish.
Notes.— This species is closely related to Cornutorogas orientalis Chen & Belokobylskij, but the new species differs by the larger eyes and subtriangular head of the female, the larger anterior tentorial pits (fig. 49), the absence of a distinct convexity of the pronotum medio-posteriorly figs 51, 53), the about 25 white segments of the antenna of the female and the largely orange-red mesoscutum.
Cornutorogas hatinhensis Long & van Achterberg, spec. nov.
Material.— Holotype, ♂ (IEBR), “Dory. 367, Vietnam: Ha Tinh (Huong Son), 1250 m, forest, 14.v.1998, K. D. Long”. Paratype: 1 ♂ (RMNH), “S. Vietnam: Dak Lak, Chu Yang Sin N.P., n[ea]r dam, 740-940 m, 1-10.vi.2007, Mal. traps, C. v. Achterberg & R. de Vries, RMNH’07”.
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Figs 18-22, Cornutorogas hatinhensis |
Holotype, ♂, length of body 3.2 mm, length of fore wing 3.3 mm.
Head.— Antenna with 49 segments, third segment 1.3 times as long as fourth segment, penultimate segment 0.6 times as long as apical segment; in lateral view scapus longer dorsally than ventrally (7:4); scapus 1.75 times longer than apically high; occipital carina strongly developed (fig. 18); eyes glabrous, emarginate; width of face equal to height of face and clypeus combined; distance between antennal sockets 0.75 times diameter of antennal socket; eye 1.9 times as long as wide; in lateral view eye 3 times as long as width of temple; POL:Od:OOL = 3:2:3 (fig. 18); face, vertex, temple and occiput shiny and smooth.
Mesosoma.— Length of mesosoma 1.45 times (without propodeal horn) its height; pronope shallow; pronotum small in dorsal view and in dorsal-lateral view hardly convex medio-posteriorly (figs 19, 20); precoxal sulcus rather deep with carinae (fig. 21); mesopleuron with dense fine punctures dorsally, and highly polished ventrally; metapleuron with sparse fine punctures; mesoscutum shiny and with sparse setae; notauli wide, deep and strongly carinate; median lobe of mesoscutum with apical carinate groove; scutellum smooth as apical part of mesoscutum; propodeum irregularly rugose-areolate with a very short carina basally; lateral horn of propodeum 0.7 times as long as long as first metasomal tergite.
Wings.— Fore wing: pterostigma 5.4 times as long as wide; length of veins r:2-SR:3-SR:SR1 = 5:11:10:34; vein r arising just before middle of pterostigma; second submarginal cell 1.4 times as long as its basal width and 1.75 times as long as its apical width; length of veins 1-CU1:cu-a:2-CU1:3-CU1 = 4:5:19:4. Hind wing: M+CU:1-M:1r-m = 20:22:7.
Legs.— Hind coxa more or less punctate; length of fore femur:tibia:tarsus:basitarsus = 30:34:34:13; length of hind femur:tibia:tarsus:basitarsus = 47:55:54:24; inner hind tibial spur about 0.3 times as long as basitarsus (7:24).
Metasoma.— Length of first tergite 1.1 times as long as its apical width, its surface very coarsely longitudinally rugose-striate; dorsal carina complete; medio-basal triangular area rather large, dorsal carina complete and ending at apex; medially second tergite 1.5 times as long as third tergite; third and fourth tergites coarsely longitudinally rugose-striate, between apical striae punctate; fifth tergite rugose-striate and mixed with punctures (fig. 22).
Colour.— Brown; head, mesoscutum dorsally, propodeum and all tergites dark brown; pronotum, fore and middle legs, mesopleuron, propodeal horns, hind coxa, trochanters and basal half of femur yellow; apical half of hind femur brown; hind tibia brown but whitish yellow basally; hind basitarsus basally, tarsus 2-4 whitish yellow; telotarsus infuscate; ovipositor sheath brown.
Notes.— This species is closely related to Cornutorogas maetoi van Achterberg, 2004, but the new species differs by the following features: pronotum slender in dorsal view, mesoscutum and scutellum smooth and fifth tergite coarsely striate, inter-mingled with discrete punctures. It differs from C. similis spec. nov. as follows: antenna of ♂ without white segments, entirely brown; mesopleuron polished and smooth ventrally, rugose-areolate dorsally, finely punctate medially; precoxal sulcus rather deep and strongly carinate (fig. 21) and propodeum irregularly rugose.
Cornutorogas hoabinhicus Long & van Achterberg, spec. nov.
Material.— Holotype, ♀ (IEBR), “Dory. 283, Vietnam: Hoa Binh (Yen Thuy), forest, 1.iii.2004, K.D. Long”.
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Figs 23-29, Cornutorogas hoabinhicus Long & van Achterberg, spec. nov., holotype, ♀. 23, Head, dorsal aspect; 24, head, anterior aspect; 25, pronotum, dorsal aspect; 26, pronotum, lateral aspect, 27, propodeum; 28, precoxal sulcus; 29, fifth tergite, dorsal aspect. Scale-line: 1 mm. |
Holotype, ♀, length of body 6.0 mm, length of fore wing 4.8 mm.
Head.— Antenna with 63 segments; penultimate segment shorter than apical segment (5:12); scapus longer dorsally than ventrally in lateral view (12:7); scapus 1.7 times longer than apical width; occipital carina strongly developed (fig. 23); face with rather distinct keel medially (fig. 16); eye largely glabrous, emarginate; in anterior view width of face equal to height of face and clypeus combined; distance between antennal sockets 0.5 times diameter of antennal socket; tentorial pits small, distance between tentorial pits twice as long as distance from pit to eye; height of eye twice as long as wide; in lateral view eye about 3 times longer than temple; POL:OD:OOL = 2:3:6 (fig. 23); face smooth with sparse fine punctures and setae; frons and area between antennal sockets with some sublongitudinal rugae (fig. 24); vertex, temple and occiput smooth and shiny.
Mesosoma.— Length of mesosoma (without propodeal horn) 1.5 times its height; pronotum long and robust in dorso-lateral view and medio-posterior area rather long and large, 0.6 times as long as pronotum (figs 25, 26); mesoscutum punctate, median lobe of mesoscutum with a smooth, wide and deep median groove posteriorly; notauli complete and strongly carinate; precoxal sulcus shallow with 5 faint carinae (fig. 28), rugose dorsally; mesopleuron smooth; scutellar sulcus wide; scutellum smooth except sparse fine punctures; propodeum coarsely longitudinal-rugose with short carina basally; propodeal spiracles round (fig. 27); in lateral view, length of propodeal horn 0.7 times as long as first tergite.
Wings.— Fore wing: pterostigma 4.7 times as long as wide; vein 1-M of fore wing distinctly curved basally; length of vein r:2-SR:3-SR:SR1 = 18:16:17:48; vein r arising from middle of pterostigma; length of veins 1-CU1:cu-a:2-CU1:3-CU1 = 5:6:35:7; second submarginal cell long, distinctly narrowed distally, its length 1.8 times as long as its basal width and 2.3 times as long as its apical width. Hind wing: M+CU:1-M:1r-m = 31:38:14.
Legs.— Hind coxa with sparse fine punctures; length of fore femur:tibia:tarsus:ba-sitarsus = 53:62:68:28; length of hind femur:tibia:tarsus:basitarsus = 86:100:105:48; inner hind tibial spur 0.2 times as long as hind basitarsus (11:48); claw with obtuse lobe.
Metasoma.— First tergite as long as its apical width, its surface very coarsely longitudinally rugose-striate; second tergite 1.2 times as long as third tergite; second and third tergites with complete carina; median length of second tergite about 1.2 times as long as third tergite (25:21); second-fourth tergites coarsely longitudinally rugose-striate, basal half of fourth tergite smooth; between striate at apex of third and fourth tergites punctulate; fifth tergite striate-punctulate (fig. 29); ovipositor sheath 0.3 times as long as hind basitarsus.
Colour.— Head, mesoscutum and mesopleuron, third-fifth tergites medio-dorsally black; basal eight segments of antenna yellow, 9th-25th segments dark brown, 28th-38th segments white and apical remainder dark brown; palpi whitish yellow; pronotum, propodeal horns, fore leg yellow (but third and fourth tarsal segments white) yellow; propodeum, first-second tergites medio-dorsally, third-fifth tergites laterally and hind coxa reddish yellow; first-second tergites laterally dark brown; fore basitarsus, second tarsal segment and telotarsus brown; middle leg yellow, but telotarsus brown; hind femur and trochantellus dark brown, basal half of hind femur reddish yellow and its apical half dark brown; basal one third of hind tibia whitish yellow, reminder dark brown; hind tarsus white, but basitarsus medially and telotarsus apically dark brown; ovipositor sheath brown.
Notes.— This species is closely related to Cornutorogas orientalis Chen & Belokobylskij, 2004, but differs by the shape of the pronotum and the shallow and sparsely carinate precoxal sulcus.
Cornutorogas similis Long, spec. nov.
Material.— Holotype, ♀ (IEBR), “Rog. 507, Vietnam: Thua Thien-Hue, Bach Ma National Park, bushes, 18.v.2007, K.D. Long”. Paratype: 1 ♂ (RMNH), same data as holotype, Rog. 508”.
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Figs 30-37, Cornutorogas similis Long, spec. nov., holotype, ♀. 30, Head, dorsal aspect; 31, pronotum, lateral aspect; 32, head, anterior aspect; 33, pronotum, dorsal aspect; 34, apical half of hind tarsus; 35, precoxal sulcus; 36, propodeum, dorsal aspect; 37, first-fifth metasomal tergites, dorsal aspect. Scale-line: 1 mm. |
Holotype, ♀, length of body 5.8 mm, length of fore wing 4.6 mm.
Head.— Antenna with 63 segments; middle segments longer than wide (6:3), penultimate antennal segment shorter than apical segment (3:4); scapus longer dorsally than ventrally in lateral view (11:7); scapus distinctly longer than apically wide (11:7); occipital carina strongly developed (fig. 32); eye largely glabrous, emarginate; in anterior view eye more than 1.6 times as high as wide (21:13); distance between antennal sockets 1.75 times diameter of antennal socket; tentorial pits rather large, distance between tentorial pits 1.25 times as long as distance from pit to eye (fig. 24); face slightly wider than height of face and clypeus combined; basal width of mandible less than lenght of malar space (6:8); in lateral view width of eye 2.4 times width of temple; POL:OD:OOL = 3:3:5 (fig. 30); face coriaceous and with sparse long setae; vertex, temple and occiput smooth and shiny.
Mesosoma.— Length of mesosoma (without propodeal horn) about 1.3 times its height; pronotum short and large in dorsal view (fig. 33) and in dorsal-lateral view hardly convex medio-posteriorly (fig. 31); side of pronotum slender (fig. 31); precoxal sulcus rather deep and strongly carinate; upper part of mesopleuron with dense fine punctures, and lower part of mesopleuron highly polished and with sparse fine punctures; mesoscutum with sparse fine punctures; notauli wide, deep and strongly carinate; median lobe of mesoscutum with a carinate long and deep groove posteriorly; scutellum with sparse fine punctures; propodeum irregularly rugose-areolate; lateral horn of propodeum 0.9 times as long as first metasomal tergite (fig. 36).
Wings.— Fore wing: pterostigma 4 times as long as wide; vein r from middle of pterostigma; length of vein r:2-SR:3-SR:SR1=7:13:12:45; length of vein 1-CU1:cu-a:2-CU1:3-CU1= 4:6:31:6; second submarginal cell long, distinctly narrowed distally, its length 1.8 times as long as apical width. Hind wing: M+CU:1-M: 1r-m = 24:29:11.
Legs.— Hind coxa with sparse fine punctures; length of fore femur:tibia:tarsus:ba-sitarsus = 53:63:58:24; length of hind femur:tibia:tarsus:basitarsus = 80:90:106:40; inner hind tibial spur 0.3 times as long as basitarsus (12:40).
Metasoma.— First tergite as long as wide apically, its surface very coarsely longitudinally rugose-striate; dorsal carinae remain separated medially (fig. 37); second tergite with very small medio-basal area and complete median carina continuing up to 0.7 of third tergite; second tergite 1.1 times as long as third tergite; second to fourth tergites coarsely longitudinally rugose-striate, and between striae punctate and punctate apically; fifth tergite with discrete punctures; length of ovipositor sheath 0.9 times hind basitarsus.
Colour.— Head, mesoscutum and mesopleuron blackish brown; pronotum white; basal ten antennal segments yellow; 11th-25th and 40th-63rd antennal segments blackish brown; 26th-39th antennal segments white; scutellum and propodeum dirty reddish brown; propodeal horns whitish yellow; sternites white, all tergites dark brown but laterally reddish yellow; fore leg white, but basitarsus and telotarsus brown; middle coxa, trochanter, tibia and second-fourth tarsal segments white; middle femur reddish yellow but dark brown apically; middle basitarsus and telotarsus dark brown; hind coxa reddish yellow but dark brown apically; hind trochantellus dark brown; apical two thirds of hind femur black and reddish yellow basally; hind tibia and basitarsus black, but basal third white; second-fourth hind tarsal segments and basal half of telotarsus white (fig. 34); wing veins and pterostigma brown; parastigma yellow.
Male.— Similar to female but antenna with 65 segments and 28th-38th antennal segments white; length of body 6.7 mm and length of fore wing 5.9 mm.
Notes.— This species is closely related to Cornutorogas orientalis Chen & Belokobylskij, but it differs from the latter by the yellow basal antennal segments, the more slender pronotum, the distinctly punctate fourth tergite, the largely smooth metapleuron and the irregularly rugose (and apically rugose-areolate) propodeum.
Cornutorogas orientalis Chen & Belokobylskij, 2004
This species has been found in North Vietnam (IEBR, RMNH: Ha Giang, Hoa Binh, Ninh Binh and Ha Tinh provinces).